This stage in our knowledge was marked by the institution of the class Cycadofilices by Potonie in 1897. Nothing, however, was known of the organs of reproduction of the Cycadofilices, until F.W. Oliver, in 1903, identified a fossil seed, Lagenostoma Lomaxi, as belonging to Lyginodendron, the identification depending, in the first instance, on the recognition of an identical form of gland, of very characteristic structure, on the vegetative organs of Lyginodendron and on the cupule enveloping the seed.
"Darwin and Modern Science"
A.C. Seward and Others